What happened: Gwyn earned the center job after taking extensive first-team reps over the past two weeks. His promotion followed a summer-long competition.
Why it matters: The opener will test a new center-quarterback exchange at regular-season speed. The assignment is a Week 1 decision rather than a permanent one.
By the numbers: Gwyn enters his fourth NFL season without a career start.
What to watch: Protection calls and interior-line communication will be the focus Sunday.